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Pyramid Scales are smaller entities deployed by the Pyramids. They have been observed on Io, where they inject Darkness into regions of Echo Mesa to undo the Traveler's work.
Overview
Pyramid Scales are smaller Pyramids were first sighted over the Echo Mesa on Io, empowering the enemies of humanity that flock towards them and reap of their paracausal power. Pyramid Scales also begin the Contact Activity, in which Guardians must fight to contain their Darkness in Gambit fashion; by killing the enemies that drop Motes of Darkness, banking them back to the Pyramid Scale and ridding any Taken Blockers sent by Quria, Blade Transform.
Trivia
The first Pyramid Scale was seen in the Garden of Salvation Raid, right after killing the Sanctified Mind, Sol Inherent, the remnants of the Pyramid it protects seemingly flies up and reveals a Scale.
Pyramid Scales seem to be "dropships" for the Darkness enemy type, similar to Skiffs for Fallen enemies and Tombships for Hive Enemies. This implies the existence of some sort of "infantry" forces for the Darkness.